2−2 Harnischia group
return to 2 Cironominae
Harnischia group is distinguished from Chironomini by the following points.
Harnischia group |
Chironomini | |
mandible | Ssome of them apical tooth pale yellow, dorsal tooth lacking |
Apical tooth and lateral teeth dark brown , with drosal tooth |
ST | simple , rarely divided into several teeth | plumose or comb shape |
SU | simple, thin or thick | plumose or simple |
mentum | Teeth arranged low convex or shallow concave, form and color various | Teeth arranged concave, mediantooth and lateral teeth dark brown. |
antenna | antenna with 5〜8 segments | antenna with 5 segments , rarely 6 segments. |
Harnischia group
Chernovskiia Cladopelma Cryptochironomus Cyphomella Demicryptochironomus Hanochironomus Harnischia Kloosia Microchironomus Paracladopelma Parachironomus Robackia Saetheria
Harnischia group is divided into four groups by form of mentum .
Robackia group |
・mentum with even teeth |
Robackia |
Chernovskiia group |
・mentum with odd teeth ・lateral teeth arranged by shallow V shape. |
Chernovskiia Harnischia Cryptochironomus Demicryptochiropnomus Kloosia |
Cyphomella group |
・mentum with odd teeth ・lateral teeth arranged by convex ・median tooth wider than 3 times of 1st lateral tooth. ・median tooth pale yellow |
Cyphomella Saetheria Paracladopelma |
Cladopelma group |
・mentum with odd teeth ・ lateral teeth arranged by convex ・median tooth not wider than 3 times of 1st lateral tooth. ・median tooth brown |
Cladopelma Microchironomus Parachironomus Saetheria Hanochironomus |
Robackia Larva body length 8〜10mm, with s.even anal setae . They were found in Kiso-river and Kumano-river. |
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Head thin, Head ratio. 0.6. Eye spots 2 pairs, slightly away. | Anal tubules :both 2 pairs long and slender , half of posterior parapod. . Posterior parapods elongate. |
Mentum with 12 teeth , almost the same size. Ventromental plate with irregular
7〜8 scales , 3rd scale largest. |
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Antenna 0.7 times of head length, 7 segments, 6th and 7th segment very
short. The blade at the top of 2nd segment reaches to the 5th segment. Two
thin and short projections at top of the 3rd the segment , but Lauterborn's
organ absent. A projection with 2 segments at the top of the 5th segment is 2 times length of the 5th, mached to 6th segment , |
Mandible: with long apical tooth and 3 inner teeth. Seta subdentalis
long, beyond the 1st inner tooth. |
STshort, SUlong needle shape. Pecten epipharyngis divided into two. Premandible with 4 teeth and one projection on back. |
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1: antenna 2: mentum 3: mandible 4:labro-epiphryngeal region 5: maxillaly palp 6: claws of anterior parapod 7: claws of posterior parapod 8:procercus and anal setae 9: head 10: anal tubules and posterior parapod |
mentum | ventromental plate | antenna | premandible | ||
Chernovskiia | Teeth are located in a line with shallow concave. The whole mentum is pale yellow. |
Fan shape , on both sides of mentum | 8 segments | 3 teeth |
A larva's forward end and back end are thin, and chest and abdomen are
thick. Posterior parapods thin, elongate backward. Maxilla long. |
Harnischia | Median tooth broad, higher than lateral teeth, brown. | Fan shape, a part of ventromental plate overlaps with mentum | 5 segments. Blade arising in the middle of segment 2. |
5〜6 | |
Cryptochironomus | Median tooth broad, pale yellow, lower than lateral teeth,. Lateral teeth dark brown. | Fan shape, outside end sharp. | 5 segments. Segment 4 and 5 very short. |
5〜6 | Ventromental plate of Demicryptochironomus more elongate than one of Cryptochironomus |
Demicryptochironomus | Median tooth broad, pale yellow, lower than lateral teeth,. Lateral teeth dark brown. | Fan shape, outside end sharp. | 7 segments.、 Blade arising on apex of segment 3. |
3 | |
Kloosia | Median tooth sharp , larger than leteral teeth. | Oval, situate both side of mentum | 6 segments. Segment 4 shorter than segment 5. |
3 | Shape of ventromental plate characteristic . |
Chernovskiia These larvae were found in sandy riverbed of 2m depth in River Kiso with following Robackia. |
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Head very small, Maxilla palup long, looks antenna. Claws of anterior parapod strong and thick. | Anal tubules elongate back-ward, 3/4 times of posterior parapod..Posterior
parapods thin ,elongate back-ward. with 10 small claws |
Median tooth and 4 pairs lateral teeth arranged by shallow V-shape .
Teeth are pale yellow all. Ventromental plate fun shape. |
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Mandible with apical tooth only, inner tooth lacking. Seta subdentalis sharp. | ST: short needle shape. SU: very long and thick. Pecten epipharyngis 4 small lobes. Premandible divided into 3 . |
Antenna 8 segments. the end in segments are indistinct. Basal segment
with a seta in center. 2 Lauterborn organs organ situate on apex of segment 4. A long oval projection on apex of segment 5. |
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Body length 7〜8mm. Head 1/38〜1/40 of body. Abdomen and chest thick. segment 1 and 11 thin.. The whole body is spindle shaped. Anal setae 8, Proceucus flat. |
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Harnischia Body lenth 6〜7mm. Body pale orange , head pale yellow. Anal setae 8. These larvae found in Lake Biwa . |
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Mediantooth with a gap on center and both side. Lateral teeth 6 pairs. Ventromental plate fan -shape with clear radial lines . |
STshort and thin, SUstrong and thick. Premandible divided into 5〜6 diatally. |
Antenna with 5 segments. Blade arising at middle of sebment 2 reaches
the point of segment 4 Lauterborn organ lacking. |
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Apical tooth of mandible sharp, 2 inner teeth small |
Cryptochironomus Body pale orange i. Larvae live in various water areas. 4 species found in Japan. |
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Median tooth pale yellow, wide, lateral teeth 5 pairs, arranged by shallow V-shape . Ventromental plate fan shape, elongate to outside. | Mandible with 2 inner teeth. | Antenna 5 segments, 2 blades( 1 short and 1 long) situated at apex of segment 2. Lauterborn organ lacking. |
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8:Anal setae 8. procercus short . 9:Anal tubules 2 pairs, conical. |
ST: thin ,long needle shape. SU: strong , long. Pecten epipharyngis triangular with small serrted teeth. Premandible divided into 5〜6 . |
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Demicryptochironomus Body length 5〜8mm.、pale orange. They live in clear water area relatively. |
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Maxilla pulp long, looks to be antenna. | Mentum : mediantooth and 7 pairs lateral teeth arrenged in shalow concave. Ventromental plate fan shape, elongate to utside. Mandible with sharp apical tooth and 2 inner teeth, dorsal tooth lacking. |
Antenna with seven segments, blade at apex of segment 3, small Lauterborn's rgan at apex of segment 5. |
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STvery thin and short, SUvery thick and long. Pecten epipharyngis triangular, with 3 teeth. Premandible with 3 large teeth. |
Anal tubules 2 pairs., conical. Procercus short, with 6〜8 anal setae |
Kloosia Body length 5〜6mm, pale orange. Whole body thin and long. Following pictuers are the larvae found in Ishigaki Iland (Okinawa pref. Japan). Other larvae found in Ina-river(Osaka :Japan) and Nabari-river ( Mie pref. Japan) have a mentum with different shape from the former. |
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Medical tooth sharp, large. with 6 pairs lateral teeth. The whole mentum
arranged in shallow V- shape. Ventromental plate situated both outside of mentum. |
Antenne with 6 segments. Blade arising at middle of sebment 2 reaches
the point of antenna. Lauterborn organ on apex of segment 3. Segment 4 shorter than segment 5. |
Mandible with 2 inner teeth. Seta subdentalis reach 2nd inner tooth |
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ST, SU: thick, long needle like together. Labral lamella triangular 2 lobes Premandible divided into 3. |
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Kloosia koreana : caught in Iahigaki-Iland. | Anal setae 8, Posterior parapod 3 times of anal tubules |
mentum | antenna | mandible | |
Cyphomella | lateral teeth 7 pairs | 5 segments, segment 3〜5 short | apical tooth and inner teeth nomal |
Saetheria(part) | lateral teeth 7 pairs | 6 segments , segment 3 long | apical tooth and inner teeth nomal |
Paracladopelma | lateral teeth 6 pairs | 5 segments, blade on halfway of segment 2、 | apical tooth thin and long, inner teeth long |
Cyphomella Body length 5〜6mm. pale orange . These larvae found in Biei-River (Hokkaido, Japan). They live in algae of surface of stone. |
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Medical tooth wide., pale yellow. Lateral teeth 7 pairs. Ventromental plate wide fan shape. radial line unirregular. |
Antenna with 5 segments, segment 3〜5 very short. 2 blades on apex of segment 2, Lauterborn's organ lacking. | ST: short. SUstrong.: Pecten epipharyngis hart shape, divided into 3 Premandible with 4 teeth. brush present. |
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Saetheria sp. Body length 5〜6mm. pale yellowish brown . Body redish orange. Anal tubules both 2 pairs long conical. Procercus short, with 7 anal setae |
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Median tooth is the size of 4 lateral teeth, pale yellow. Lateral teeth
7 pairs. Ventromental plate wide fan shape, radial line very clear. |
Antenna with 6 segments, segment 4〜6 short A blade at halfway of segment 2 beyond tip of antenna. Lauterborn's organ on apex of segment 3 short, with baloon shape projection. |
ST: short. SU: strong, long needle like. Pecten epipharyngis 1 blade, triangular. Premandible with 4 teeth. |
Paracladopelma Body length 5^6mm. Head pale yellow, body pale orange. 3 species found. in Japan. |
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Median tooth arc , pale yellow. Lateral teeth 6 pairs; expect 1st.lateral tooth brown, 1 st. paleyellow. Ventromental plate wide fan shape, radial lines about 20 clear. | ST: very thin, short needle like. SU: long needle like. Pecten epipharyngis small triangular. Premandible with 4 teeth, brush lacking. |
Apical tooth long, sharp. with 2 pointed inner teeth Seta subdentalis long needle shape. Pecten mandibularis consisting of 2. |
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Antenna with 5 segments. Blade arising downward 2/3 of segment 2 reaches
apex of antenna. Segment 3〜5 short. Lauterborn's organ lacking. |
Eye spot 2 pairs , small. Anal tubules long conical. Procercus short, with 8 anal setae. |
mentum | ventromantal plate | antenna segment |
pecten epipharyngis | pramandible | |
Cladopelma | Center of Median tooth with a gap . Outer 3 lateral teeth characteristic. |
nomal: fan shape | 5 | divited into 3 lobes | distally bifid |
Microchironomus | Median 3 teeth arranged in a mound shape.、 Outer 3 lateral teeth characteristic. |
nomal: fan shape | 5 | simple, with serrated edge | distally bifid |
Parachironomus | Nomal: 1 median tooth and 7 pairs lateral teeth. | small fan shape、 upperside edge rough |
5 | triangular with rough serratered both sides | distally bifid |
Saetheria(hirta) | Median tooth 2 times of 1st. lateral tooth. | large fan shape | 6 | 2 lobes triangular | distally divited into 4〜5 |
Saetheria(tylus) | Median tooth 4 times of 1st. lateral tooth. | large fan shape | 5 | 2 lobes triangular | distally divited into 4 |
Hanochironomus | Median 3 teeth arranged in a mound shape. | large fan shape | 6? | details unclear | details unclear |
Cladopelma Body length 5〜6 mm. pale orange. Head yellowish brown, under side blackish brown. This larva was found in Lake-BIwa only. |
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Mentum with 15 teeth,. Median tooth with a small gap in center. 1st〜4th
lateral teeth the same size, outer 3 teeth arranged mound shape. Ventromental plate fan shape, radial line clear. |
ST. SUthick and long both. Pecten epipharungis 3 triangular lobes. Premandible distally bifid . |
Antenna with 5 segments, Lauterborn organ small. |
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2 inner teeth distally flat. |
Microchironomus Body length 5〜6mm, Head yellowish brown, body pale orange . They found in Lake Biwa and River Nagara. |
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Mentum with 13 teeth, 3 median teeth arranged in mound like. 1st 〜 3 lateral teeth the same size. 4th 〜6 lateral teeth arranged in mound like. Ventromental plte wide fan shape., radial line clear. | 2 inner teeth distally flat Seta subdentalis spatula like. |
ST, SUthick needle like. Pecten epipharyngis heart shape , distally serrated Premandible divided into 2. |
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anal tubules 2 pairs ,spindle shape. Procercus : height and diameter the same size. , anal setae 8 |
Parachironomus Body length 5〜6mm. Head pale brown, body red. 2 species found. |
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Mentum with 15 teeth, median tooth larger than lateral teeth. All
of the teeth pale brown and sharp. Ventromental plate small fan shape、 upper edge rough. radial line clear, rough. |
ST,SUboth thick and long needle like. Pecten epipharyngis heart shape, side edge with several rough serrated Premandible divided into 2. |
Antenna with 5 segments. Lauterborn rgan small. Blade reach the tip of antenna . 。 |
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・Eye spot 3. ・Mandible : a large apical tooth and 2 inner teeth. Seta subdentalis very sharp. ・Procercus short, anal setae 7. |
Saetheria
Saetheria (part) incluede following two types.
Saetheria hirta type :Body length about 6mm . Head thin , pale yellowish brown . Body
pale yellowish orange .
They live in nomal or fast flows ; sand or pebble river bed,
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Mentum with 13 teeth, mediantooth the largest of all, lateral teeth 6
pairs. Ventromental plate large fan shape, radial line clear. |
Mandible: Apical tooth large, 3 inner teeth small. Blade sharp, needle like. Anal setae 8, procercus short. |
ST: thin long needle like. SU: thick, strong. Pecten epipharyngis divided into 2 distally. Premandible with 4〜5 teeth. |
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Antenna with 6 segments. segment 4〜6 very short. Blade seated in distal 2/5 of segment 2, accessory blade 1/5 of blade. Small Lauterborn's organ seated on segment 3 distally. Style longer than Lauterborn's organ. |
Saetheria tylus type:Body length 4〜5mm. Head pale yellowish brown, body reddish orange. They found in Lake Biwa, Lake Shikotu, live in a bottom of water depth 1〜2m、and in River Hijikawa ,R.Amahuri(Kagosima pref.) |
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Median tooth of mentum wide arc, the same of 4 lateral teeth. Lateral teeth with 6 pairs , 3rd. and 6th. tooth small. Ventromental plate fan shape, with about 10〜20 lines. |
ST,SUboth thick needle like. Pecten epipharyngis 2 triangular plates. Premandible with 4 teeth, brush lacking. |
Apical tooth and 3 inner teeth both thin , pale brown. Seta subdentalis reach 2nd. inner tooth. |
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Antenna with 5 segments, 3rd.〜5th. segments short. Lauterborn's organ lacking, long style present. Blade reach tip of antenna. |
Anal setae 7〜8, procercus short. |
not detaile datas
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